Skirmishing/Wilderness Survival:
The first two fit (different bonuses on swap).
The third 10% more damage on flanking seems better suited to replace Peak Strength in Wilderness Survival.
Replace Peak Strength with Speed Shot. On swap, gain quickness. Now you would have swiftness, fury and quickness and could really put some initial burst up when you swap.
Nature:
Replace Bountiful Hunter with Skilled Healer (All Heal Skills recharge 20% faster).
Fits with other 2 protection based skills.
Beastmastery:
Instinctual Bond: Change to Difficult Target: 50% of all non-melee attacks miss pet (hard to hit or thick skinned).
Nature’s Wraith: Change to Wild Aggression: Pet and Ranger do X% more damage for first Y seconds of a fight. (Help me define the variables).